Paleontology in Pennsylvania refers to paleontological research occurring within or conducted by people from the U.S. state of Pennsylvania. The geologic column of Pennsylvania spans from the Precambrian to Quaternary.During the early part of the Paleozoic, Pennsylvania was submerged by a warm, shallow sea. This sea would come to be inhabited by creatures like brachiopods, bryozoans, crinoids, graptolites, and trilobites. The armored fish appeared during the Silurian. By the Devonian the state was home to other kinds of fishes. On land, some of the world's oldest tetrapods left behind footprints that would later fossilize. Some of Pennsylvania's most important fossil finds were made in the state's Devonian rocks. Carboniferous Pennsylvania was a swampy environment covered by a wide variet