A leaf (plural leaves) is the principal lateral appendage of the vascular plant stem, usually borne above ground and specialized for photosynthesis. The leaves, stem, flower and fruit together form the shoot system. Leaves are collectively referred to as foliage, as in "autumn foliage". In most leaves, the primary photosynthetic tissue, the palisade mesophyll, is located on the upper side of the blade or lamina of the leaf but in some species, including the mature foliage of Eucalyptus, palisade mesophyll is present on both sides and the leaves are said to be isobilateral. Most leaves are flattened and have distinct upper (adaxial) and lower (abaxial) surfaces that differ in color, hairiness, the number of stomata (pores that intake and output gases), the amount and structure of epicuticul