Tholodus is an extinct genus of basal ichthyopterygian known from the Middle Triassic (mid-late Anisian to late Ladinian stage) of Germany, northeastern Italy and possibly China. It was first named by Christian Erich Hermann von Meyer in 1851 and the type species is Tholodus schmidi. It is known from many disarticulated and fragmentary remains, mainly teeth and jaw fragments. Most specimens were collected from various localities across the Ladinian-aged Muschelkalk, Germany, mainly from the Jena Formation of the upper Lower Muschelkalk, where the holotype was found. Dalla Vecchia (2004) recently described two additional specimens, a mandibular ramus and a maxilla, both bearing teeth and nearly uncrushed, and some postcranial remains, from a single late Anisian outcrop, from the southern Al