The branching system of Chara species is complex with branches derived from apical cells which cut off segments at the base to form nodal and internodal cells alternately. The main axes bear whorls of branches in a superficial resemblance to Equisetum. They are typically anchored to the littoral substrate by means of branching underground rhizoids. Chara plants are rough to the touch because of deposited calcium salts on the cell wall. The metabolic processes associated with this deposition often give Chara plants a distinctive and unpleasant smell of hydrogen sulfide.