Lycaenidae is the second-largest family of butterflies (behind Nymphalidae, brush-footed butterflies), with over 5,000 species worldwide, whose members are also called gossamer-winged butterflies. They constitute about 30% of the known butterfly species. The family is traditionally divided into the subfamilies of the blues (Polyommatinae), the coppers (Lycaeninae), the hairstreaks (Theclinae) and the harvesters (Miletinae); others include the Lipteninae, Liphyrinae, Curetinae and Poritiinae, but some authorities also include the family Riodinidae within the Lycaenidae.